ARTHROPODA
Introduction
NUMERICAL STRENGTH

HABITS AND HABITATS

SIGNIFICANCE OF STUDY

HISTORY

DERIVATION OF NAME

DEFINITION

GENERAL CHARACTERS

CLASSIFICATION

Class 1
Crustacea
(Larval forms of crustacea)


DISTINCTIVE CHARACTERS OF CRUSTACEA

OTHER CRUSTACEA
 

TRILOBITA
Distribution
Habbits and habbitat
Structure
Development
Position of Trilobites

ARACHINDA
DISTRIBUTION
HABITS AND HABITATS
EXTERNAL FEATURES
BODY WALL
BODY CAVITY AND MUSCULATURE
ENDOSKELETON
DIGESTIVE SYSTEM
CIRCULATORY SYSTEM
RESPIRATORY SYSTEM
EXCRETORY SYSTEM NERVOUS SYSTEM
SENSE ORGANS
REPRODUCTIVE SYSTEM
LIFE HISTORY

DISTINCTIVE CHARACTERS OF ARACHINDS

DOUBTFUL ARACHINDS

MYRIAPODA
DISTINCTIVE CHARACTERISTICS
CLASSIFICATION OF MYRIAPODA
MILLIPEDES
CENTIPEDES

ONYCHOPHORA
DISTRIBUTION
HABITS AND HABITATS
EXTERNAL FEATURES
INTERNAL ANATOMY
AFFINITIES AND PERIPATUS

INSECTA
(THE COMMON COCKROACH)
Periplaneta americana

DISTRIBUTION
HABITS AND HABITAT
EXTERNAL FEATURES
BODY CAVITY
DIGESTIVE SYSTEM
BLOOD VASCULAR SYSTEM
RESPIRATORY SYSTEM
EXCRETORY SYSTEM
NERVOUS SYSTEM
SENSE ORGANS
REPRODUCTIVE SYSTEM
LIFE HISTORY

GRASS HOPPER OR LOCUST

DISTRIBUTION
HABITS AND HABITAT
EXTERNAL FEATURES
BODY CAVITY
DIGESTIVE SYSTEM
BLOOD VASCULAR SYSTEM
RESPIRATORY SYSTEM
EXCRETORY SYSTEM
NERVOUS SYSTEM
SENSE ORGANS
REPRODUCTIVE SYSTEM
LIFE HISTORY

 

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The ‘success’ of a person in the human society is judged by his bank balance and the place he occupies in the society. But when we talk oft the ‘biological success’ of man or any other animal as a species, we consider upon quite different criteria such as:

(I)        The numbers of species and individuals.

(2)        Stretches of territory occupied. -

(3)        Variety of habitats.

(4)        Amount and kinds of food consumed.

(5)        capabilities, of defending against enemies, etc. By these standards, Phylum Arthropoda occupies the first

place in the Animal Kingdom. It is the largest and the most dominant group of animals on the earth at present.

NUMERICAL STRENGTH

 Arthropoda represents a vast assemblage of animals. No one knows how many species of arthropods there are in ‘the world.. it is the largest group in the animal kingdom, outnumbering all the rest by live to one of the total known species of the animals, more than four-fifths i.e.. one million (10,00,000) are arthropods. Of Insecta alone, which constitute only one of the six major classes of arthropods, there are more species (90,00,000) than of all the other 77 or 80 classes of the animal kingdom put together. Many of them are enormously abundant as individuals. However, speaking of biological success in the terms of numbers of species and numbers of individuals, the arthropods have some very close rivals in the Phylum Nematoda. When all the round-worms are finally catalogued, they may eventually outnumber even the arthropods.

HABITS AND HABITATS

 They have most successfully established themselves all over the globe in diverse habitats, wherever life is possible on earth. They occur over high mountain ranges, 20,000 feet above the sea level, and have been encountered by airplanes flying miles above the earth. They have been brought up from the deepest sea bottoms that have been dredged, some obtained from depths more than five miles in the oceans. They inhabit fresh-water lakes, streams, ponds, sulphur springs, hot springs and even pools of petroleum. They live in. the under desert sends,  as parasites in and on the body of other animals and as pests on plants. They have even invaded the air where they are the most successful of animals. They fly, hop, crawl and just sit still. Some are gregarious,
while colonial insects have evolved social organizations with a well-marked division of labour among members of different castes.

 

SIGNIFICANCE OF STUDY 

Arthropods are economically of great importance to man while men are feverishly contending with each other for materialistic success in terms of fame and fortune; myriads of arthropods are quietly but effectively competing with humans for supremacy in the Animal World. Man has had a very hard time living with the arthropods because of the billions of rupees worth of damage they cause to his economy each year. They consume the largest amount and kinds of food and are most capable of defending themselves against their enemies. There is probably no species of organism that is not on occasion eaten by some arthropod. In turn, the arthropods are eaten by many other animals.

They play an important role in the human welfare and health. Strangely enough man can hardly live Without them. Some arthropods are beneficial to man while others are his dreadful enemies. On one hand, many members, such as large crabs, shrimps, prawns, lobsters and locusts, etc., contribute to man’s food supply; others pollinate flowers, destroy pests, supply useful products and provide other valuable services. On the other hand, they destroy crops, food-stores, house­hold goods, clothing, furniture, etc., inflict injury as parasites and transport organisms of dreadful diseases to man, to his livestock and other creatures. . Many fertile regions of the world are still closed to man on account of the presence of the disease-bearing arthropods. The insects, in particular, are the man’s chief competitors for food and shelter. Several times in the history of man, insects, the most successful of all arthropods, have threatened to wipe out man’s food supply and destroy him with disease.

Modern man, with his growing knowledge of Zoology and Chemistry, is gradually learning to cope with his arthropod rivals but the strange paradox creates serious problem in arthropod management; for, if we were to destroy all of them, because many were injurious, we would at the same time destroy the beneficial varieties. Consequently, man has got to live with these extremely prolific and efficient creatures and share some of his with them. 

HISTORY

The arthropods belong to an ancient stock and were well diversified in the past. Today, they are the most dominant animals on the earth, if numbers of different species are accepted as criteria of dominance.

Aristotle first described crabs and some other Decapoda and named them Malacostraca or the soft-bodied animals.

 The great taxonomist Linnaeus put Malacostraca under Insecta which was a very heterogeneous group exactly equivalent to the modern Arthropoda. Linnaeus made the group Insecta opt era to receive the crustaceans, spiders and the myriapods.

 Jean Baptiste De Monet, Chevalier De Lamarck, the great French philosophical botanist, divided the Linnean Insecta into three crasses— Crustacea, Hexapoda and Arachnida. The name Crustacea was created by Cuvier in 1800, and Lamarck put lobsters, crabs, water-fleas etc., in this class. The name Hexapoda was created for the modern insects but it has not been universally adopted in favour of the present name insecta. Lamarck composed Arachnida for scorpions, spiders, mites, Myriapoda and Thysanura. The name Myriapoda was created by Latreille in l796. The Thysanura are now known to be primitive insects, while Myriapoda is an independent class.

 Cuvier placed all these animals together with Annelida under the group Articulata. In 1845, C. T. E. Von Siebold separated Annelida from Articulata to make a class of them under the Vermes, while he combined Crustacea, Arachnida and Hexapoda under an independent group, to which he gave the name Arthropoda.

 DERIVATION OF NAME

The name Arthropoda means ‘jointed legs’, and refers to the most characteristic structure of them. It is, therefore, the jointed nature of the limbs, which has earned the name Arthropoda (Gk., Arthros=jointed; podos= foot) for this phylum.

 DEFINITION

 The Arthropoda in general are metamerically segmented coelomate and jointed-limbed animals, with a thick exoskeleton containing chitin, moulting periodically, and without true nephridia and cilia in their structure.

 GENERAL CHARACTERS

It is well presumed that the two phyla had a common origin about 500 million years ago.

  • (1)        The body is bilaterally symmetrical and metamerically segmented, i.e., composed of a longitudinal’ series of similar somites or segments. Some of the anterior segments undergo cephalization, forming a distinct head. Arthropods show the highest degree of cephalizat ion. of all the invertebrates with the possible exception of cephalopods.
  • (2)        Externally, the body is covered with a thick, tough and non-living, chitinous cuticle, forming the exoskeleton. This forms a protective armour around the body, furnishes a supporting framework for the softer body-parts within and provides attachment for the muscles.. The exoskeleton remains thin, soft and flexible between the segments so that movements are possible.
  • (3)        The exoskeleton is made of dead matter so that it cannot grow. As the animal grows in size with age, the old exokeleton is cast off periodically and a new one developed from the underlying epi~ dermis, slightly bigger than its predecessor to accommmodate the increased body. This phenomenon is termed moulting or ecdysis and is quite characteristic of the Arthropoda.
  •  (4)        The segments of the body usually bear paired,  lateral       and     jointed   appendages, which are variously modified as jaws, gills, legs, etc.
  •  (5)        The musculature consists of separate muscles, instead of continuous muscular layers of the body-wall of an annelida. All the
     muscles are striped.
  • (6)        The body is triploblastic. The body-cavity is a haemocoel the  true coelom is greatly reduced in the adult, represented only by the
    cavities of the reproductive and excretory organs.
  • (7)        The circulatory system is open, i.e., the capillaries are absent and the arteries open into irregular spaces, called lacunae or sinuses which collectively form the haemocoel.
  • (8)        The respiration is carried on either through general body surface, or through gills in the aquatic forms, and tracheae or book-lungs,  in the terrestrial forms.
  • (9)        True nephridia are absent. Excretion occurs by green glands  (coelomoducts) or by malpighian tubules.
  • (10)      The nervous system is of the annelidan type. The dorsal brain is  connected by a nerve-ring round the gullet with a double and  ganglionated ventral nerve cord.
  • (11)      Compound eyes with mosaic vision have developed.
  • (12)      The cilia and flagella are entirely absent in their structure and even the sperms are devoid of them
  • (13)      The sexes are generally separate, and the sexual dimorphism is usually evident. The gonads and their ducts are usually paired. Fertilization is internal and the development includes little or more metamorphosis. Parental care is often well marked ; many of them carry their eggs until they hatch, and often the young animals are also carried.

 CLASSIFICATION

Arthropoda is heterogeneous group including a variety of animals. There arc more species of arthropods known to biologists than of all other kinds of living organisms put together. Consequently, there is great disagreement regarding its classification. Some divide it into five classes while others into as many as thirteen classes. However, it seems quite convenient at present to recognize only five classes as shown below:

Class    I. Crustacea. (L., crusta = a hard shell)—Mostly aquatic. Exo­skeleton impregnated with lime salts. Two pairs of antennae and three pairs of jaws. Respiration by gills.

Examples:        Crabs, Prawns, Lobsters, Barnacles, etc.

 

Class    II. Aracbnoidea. (G., arachne =.spider. oid=like)—Air breathing, terrestrial arthropods. Antennae and true jaws absent. Four pairs of logs.

Examples:        ‘Spiders. .Scorpions, King-crabs, etc.

 

Class    Ill. Insects. (L., insectum=having been cut into)—Terrestrial. air-breathing arthropods. Body with distinct head, thorax and abdomen. One pair of antennae, three pairs of jaws, three pairs of walking legs and one or two pairs of wings. Respiration by tracheae.

Examples :       Cockroaches, Houseflies, Silkrnoih, Butterflies, Bees, Mosquitoes, Bugs, Locusts, Ants, Wasps. etc.

 

Class    IV. Myriapoda. (G., myrios = 10 thousand; poda = foot)-Air breathing arthropods with many segmented body One pair of antennae, three pairs of jaws and many pairs of legs.

Examples : Millipedes and Centipedes.

 

Class    V. Onychophora. (G., onychos= claw; phoros= bearing) —-Terrestrial, air breathing, primitive, worm-like. Single pair of antennae, eyes and jaws, Numerous stumpy legs.

Example:         Peripatus.

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